Lily's relationship with The Crown actor Matt Smith is definitely her most-publicised romance to date. The duo first on the set of Pride and Prejudice and Zombies in 2014 and sparked relationship rumours shortly after. Although they made a number of appearances together, they usually refrained from talking about each other in interviews. After five years of dating, it was reported that the couple had split in December 2019. Even though they reunited for a walk in London in March 2020, it appears that they've officially parted ways.